Compilation of selected papers published on Permian topics in 2021


Biakov A.S., Zakharov Y.D., Horacek M. & Goryachev N.A. (2021). On the position of the Wuchiapingian-Changhsingian boundary in the North-East Russia according to radioisotopic and chemostratigraphic data. Doklady Earth Sciences, 500 (2), 816-819.

Brynko I.V., Polzunenkov G.O., Biakov A.S. & Vedernikov I.L. (2022). The First U-Pb SHRIMP dating of zircons from Capitanian (Middle Permian) deposits of the Omolon Massif (Northeast Russia). Russian Journal of Pacific Geology, 15 (1), 51-59.

Dobbs S. C., Riggs N. R., Marsaglia K. M., González-León C. M., Cecil M. R. & Smith M. E. (2021). The Permian Monos Formation: Stratigraphic and detrital zircon evidence for Permian Cordilleran arc development along the southwestern margin of Laurentia (northwestern Sonora, Mexico). Geosphere, 17(2), 520-537.

González C. R. & Saravia P. D. (2021). Proposed chronostratigraphic units for the Carboniferous and Early Permian of the southwestern Gondwana margin. Geological Society, London, Special Publications, 512.

Heuer F., Leda L., Moradi-Salimi H., Gliwa J., Hairapetian V. & Korn D. (2021). The Permian–Triassic boundary section at Baghuk Mountain, Central Iran: carbonate microfacies and depositional environment. Palaeobiodiversity and Palaeoenvironments, 1-20.

Kolar-Jurkovšek T., Hrvatović H., Aljinović D., Nestell G. P., Jurkovšek B. & Skopljak F. (2021). Permian-Triassic biofacies of the Teočak section, Bosnia and Herzegovina. Global and Planetary Change, 200, 103458.

Korn D., Leda L., Heuer F., Moradi Salimi H., Farshid E., Akbari A., Schobben M., Ghaderi A., Struck U., Gliwa J., Ware D. & Hairapetian V. (2021). Baghuk Mountain (Central Iran): high-resolution stratigraphy of a continuous Central Tethyan Permian–Triassic boundary section. Fossil Record, 24(1), 171-192.

Lützner H., Tichomirowa M., Käßner A. & Gaupp R. (2021). Latest Carboniferous to Early Permian volcano-stratigraphic evolution in Central Europe: U–Pb CA–ID–TIMS ages of volcanic rocks in the Thuringian Forest Basin (Germany). International Journal of Earth Sciences, 110(1), 377-398.

Ma Q., Ye F., Wang W., Sheng Y., Shu L. & Chen X. (2021). Grid mapping revealed hidden geochemical lens and its chemostratigraphic bias in the Middle‐Upper Permian marine carbonate sequence in Laibin, South China. Geological Journal, 56(12), 6088-6101.

Shi G. R., Nutman A. P., Lee S., Jones B. G. & Bann G. R. (2022). Reassessing the chronostratigraphy and tempo of climate change in the Lower-Middle Permian of the southern Sydney Basin, Australia: Integrating evidence from U–Pb zircon geochronology and biostratigraphy. Lithos, 410, 106570.

Wainman C. C., McCabe P. J. & Crowley J. L. (2021). New insights on the age and stratigraphy of the Cisuralian succession in the Cooper Basin, Australia, based on U–Pb CA-TIMS dating of volcanic air-fall tuffs. Australian Journal of Earth Sciences, 1-12.

Yang W., Wan M., Crowley J. L., Wang J., Luo X., Tabor N., Angielczyk K. D., Gastaldo R., Geissman J., Liu F., Roopnarine P. & Sidor C. A. (2021). Paleoenvironmental and paleoclimatic evolution and cyclo-and chrono-stratigraphy of Upper Permian–Lower Triassic fluvial-lacustrine deposits in Bogda Mountains, NW China—Implications for diachronous plant evolution across the Permian–Triassic boundary. Earth-Science Reviews, 222, 103741.

Zhang L., Dong D., Qiu Z., Wu C., Zhang Q., Wang Y., Liu D., Deng Z., Zhou S. & Pan S. (2021). Sedimentology and geochemistry of Carboniferous-Permian marine-continental transitional shales in the eastern Ordos Basin, North China. Palaeogeography, Palaeoclimatology, Palaeoecology, 571, 110389.

Zhang M., Qin H. F., He K., Hou Y. F., Zheng Q. F., Deng C. L., He Y., Shen S. Z., Zhu R. X. & Pan Y. X. (2021). Magnetostratigraphy across the end-Permian mass extinction event from the Meishan sections, southeastern China. Geology, 49(11), 1289-1294.

Zhao H., Lyu Z., Chen Z. Q., Algeo T. J., Orchard M. J., Liu Y., Hu Z., Zhang L. & Zhang X. (2021). Integrated biochemostratigraphy of the Permian-Triassic boundary beds in a shallow carbonate platform setting (Yangou, South China). Global and Planetary Change, 206, 103583.

Geochemical proxies and palaeoclimate

Davydov V. I., Karasev E. V., Nurgalieva N. G., Schmitz M. D., Budnikov I. V., Biakov A. S., Kuzina D. M., Silantiev V. V., Urazaeva M. N., Zharinova V. V., Zorina S. O., Gareev B. & Vasilenko D. V. (2021). Climate and biotic evolution during the Permian-Triassic transition in the temperate Northern Hemisphere, Kuznetsk Basin, Siberia, Russia. Palaeogeography, Palaeoclimatology, Palaeoecology, 573, 110432.

Dudás F. O., Zhang H., Shen S. Z. & Bowring S. A. (2021). Major and Trace Element Geochemistry of the Permian-Triassic Boundary Section at Meishan, South China. Frontiers in Earth Science, 9.

Fang Q., Wu H., Shen S. Z., Zhang S., Yang T., Wang X. & Chen J. (2021). Trends and Rhythms in Climate Change During the Early Permian Icehouse. Paleoceanography and Paleoclimatology, 36(12), e2021PA004340.

Frank T. D., Fielding C. R., Winguth A. M. E., Savatic K., Tevyaw A., Winguth C., McLoughlin S., Vajda V., Mays C., Nicoll R., Bocking M. & Crowley J. L. (2021). Pace, magnitude, and nature of terrestrial climate change through the end-Permian extinction in southeastern Gondwana. Geology, 49(9), 1089-1095.

Grasby S. E., Bond D. P. G., Wignall P. B., Yin R., Strachan L. J. & Takahashi S. (2021). Transient Permian-Triassic euxinia in the southern Panthalassa deep ocean. Geology, 49(8), 889-893.

Hu Z., Li W., Zhang H., Krainer K., Zheng Q. F., Xia Z., Hu W. & Shen S. Z. (2021). Mg isotope evidence for restriction events within the Paleotethys ocean around the Permian-Triassic transition. Earth and Planetary Science Letters, 556, 116704.

Kaiho K., Aftabuzzaman M., Jones D. S. & Tian L. (2021). Pulsed volcanic combustion events coincident with the end-Permian terrestrial disturbance and the following global crisis. Geology, 49(3), 289-293.

Kani T. & Isozaki Y. (2021). The Capitanian minimum: a unique Sr isotope beacon of the latest Paleozoic seawater. Frontiers in Earth Science, 751.

Li Q., Azmy K., Yang S., Xu S. L., Yang D., Zhang X. H., Chen A. Q. & Chen H. D. (2021). Early–Middle Permian strontium isotope stratigraphy of marine carbonates from the northern marginal areas of South China: Controlling factors and implications. Geological Journal, 56(3), 1658-1672.

Newby S.M., Owens J.D., Schoepfer S.D. & Algeo T. J. (2021). Transient ocean oxygenation at end-Permian mass extinction onset shown by thallium isotopes. Nature Geoscience, 14, 678–683.

Onoue T., Soda K. & Isozaki Y. (2021). Development of Deep-Sea Anoxia in Panthalassa During the Lopingian (Late Permian): Insights From Redox-Sensitive Elements and Multivariate Analysis. Frontiers in Earth Science, 685.

Pfeifer L. S., Birkett B. A., Van Den Driessche J., Pochat S. & Soreghan G. S. (2021). Ice-crystal traces imply ephemeral freezing in early Permian equatorial Pangea. Geology, 49(11), 1397-1401.

Takahashi S., Hori R. S., Yamakita S., Aita Y., Takemura A., Ikehara M., Xiong Y., Poulton S. W., Wignall P. B., Itai T., Campbell H. J. & Spörli B. K. (2021). Progressive development of ocean anoxia in the end-Permian pelagic Panthalassa. Global and Planetary Change, 207, 103650.

Wang W. Q., Katchinoff J. A., Garbelli C., Immenhauser A., Zheng Q. F., Zhang Y. C., Yuan D. X., Shi Y. K., Wang J., Planavsky N. & Shen S. Z. (2021). Revisiting the Permian seawater 87Sr/86Sr record: New perspectives from brachiopod proxy data and stochastic oceanic box models. Earth-Science Reviews, 218, 103679.

Wei H., Tang W., Gu H., Fu X. & Zhang X. (2021). Chemostratigraphy and pyrite morphology across the Wuchiapingian‐Changhsingian boundary in the Middle Yangtze Platform, South China. Geological Journal, 56(12), 6102-6116.

Wu W., Liu W., Mou C., Liu H., Qiao Y., Pan J., Ning S., Zhang X., Yao J. & Liu J. (2021). Organic-rich siliceous rocks in the upper Permian Dalong Formation (NW middle Yangtze): Provenance, paleoclimate and paleoenvironment. Marine and Petroleum Geology, 123, 104728.

Zakharov Y.D., Horacek M. & Biakov A.S. (2021). Variations in Nitrogen isotope composition in clay deposits of the Permian-Triassic boundary beds in the Verkhoyansk Region (Northeast Asia) and their implication for reconstruction of marine environments. Stratigraphy and Geological Correlation, 29 (20), 192-214.

Palaeoecology and palaeoenvironments

Caracciolo L., Ravidà D. C. G., Chew D., Janßen M., Lünsdorf N. K., Heins W. A., Stephan T. & Stollhofen H. (2021). Reconstructing environmental signals across the Permian-Triassic boundary in the SE Germanic Basin: A Quantitative Provenance Analysis (QPA) approach. Global and Planetary Change, 206, 103631.

Ding Y., Duan Y., Wu Y. & Cao C. (2021). Trace fossils from the Permian Lopingian Talung Formation at the northern Penglaitan section of Laibin area, South China: Ichnology, palaeoenvironment, and palaeoecology. Geological Journal, 56(12), 6117-6134.

Lloret J., López-Gómez J., Heredia N., Martín-González F., De la Horra R., Borruel-Abadía V., Ronchi A., Barrenechea J., García-Sansegundo J., Galé C., Ubide T., Gretter N., Diez J.B., Juncal M. & Lago M. (2021). Tectono-sedimentary evolution in the Pyrenean-Cantabrian orogenic belt (N Spain): insights into the transition between the Variscan and Alpine cycles in near-equator Permian European basins. Global and Planetary Change, 207, 103677.

Lloret, J., De la Horra., R., Barrenechea, J.F., Gretter, N., López-Gómez, J. & Ronchi, A. (2021). Permian and Triassic paleosols in the fluvial-lacustrine record of the central Pyrenees Basin, Spain: A stratigraphic tool for interpreting syn-tectonic sedimentary evolution and paleoclimate. Newsletters on Stratigraphy 54(3), 377-404.

Lopes G., Pereira Z., Fernandes P., Mendes M., Marques J. & Jorge R. C. (2021). Late Permian palaeoenvironmental evolution of the Matinde Formation in the Muarádzi Sub-basin, Moatize-Minjova Basin, Mozambique. Journal of African Earth Sciences, 176, 104138.

Shcherbakov D. E., Vinn O. & Zhuravlev A. Y. (2021). Disaster microconchids from the uppermost Permian and Lower Triassic lacustrine strata of the Cis-Urals and the Tunguska and Kuznetsk basins (Russia). Geological Magazine, 158(8), 1335-1357.

Tian X. & Wang Y. (2021). Compositional characteristics and paleoenvironmental significance of Changhsingian (Late Permian) sponge reefs in South China. Arabian Journal of Geosciences, 14(12), 1-28.

Torres-Martínez M. A., Vinn O. & Martín-Aguilar L. (2021). Paleoecology of the first Devonian-like sclerobiont association on Permian brachiopods from southeastern Mexico. Acta Palaeontologica Polonica, 66(1).

Zheng Q. F., Zhang H., Yuan D. X., Wang Y., Wang W. Q., Cao C. Q. & Shen S. Z. (2021). High-resolution sedimentology, ichnology, and benthic marine redox conditions from Late Permian to the earliest Triassic at Shangsi, South China: Local, regional, and global signals and driving mechanisms. Earth-Science Reviews, 103898.

Zhu J., Zhang Z., Santosh M., Tan S., Deng Y. & Xie Q. (2021). Recycled carbon degassed from the Emeishan plume as the potential driver for the major end-Guadalupian carbon cycle perturbations. Geoscience Frontiers, 12(4), 101140.

Palaeogeography and tectonics

Dan W., Wang Q., Murphy J. B., Zhang X. Z., Xu Y. G., White W. M., Jiang Z. Q., Ou Q., Hao L. L. & Qi Y. (2021). Short duration of Early Permian Qiangtang-Panjal large igneous province: Implications for origin of the Neo-Tethys Ocean. Earth and Planetary Science Letters, 568, 117054.

Domeier M., Font E., Youbi N., Davies J., Nemkin S., Van der Voo R., Perrot M., Benabbou M., Boumehdi M. A. & Torsvik T. H. (2021). On the Early Permian shape of Pangea from paleomagnetism at its core. Gondwana Research, 90, 171-198.

Fan J. J., Niu Y., Luo A. B., Xie C. M., Hao Y. J. & Liu H. Y. (2021). Timing of the Meso-Tethys Ocean opening: evidence from Permian sedimentary provenance changes in the South Qiangtang Terrane, Tibetan Plateau. Palaeogeography, Palaeoclimatology, Palaeoecology, 567, 110265.

Guan C., Yan M., Zhang W., Zhang D., Fu Q., Yu L., Xu W., Zan J., Li B., Zhang T. & Shen M. (2021). Paleomagnetic and chronologic data bearing on the Permian/Triassic boundary position of Qamdo in the Eastern Qiantang Terrane: Implications for the closure of the Paleo‐Tethys. Geophysical Research Letters, 48(6), e2020GL092059.

Kim S. W., Kwon S., Jeong Y. J., Kee W. S., Lee B. C., Byun U. H., Ko K., Cho D. L., Hong P. S., Park S. I. & Santosh M. (2021). The Middle Permian to Triassic tectono-magmatic system in the southern Korean Peninsula. Gondwana Research, 100, 302-322.

Lawton T. F., Blakey R. C., Stockli D. F. & Liu L. (2021). Late Paleozoic (Late Mississippian–Middle Permian) sediment provenance and dispersal in western equatorial Pangea. Palaeogeography, Palaeoclimatology, Palaeoecology, 572, 110386.

Niu Y. Z., Shi G. R., Ji W. H., Zhou J. L., Wang J. Q., Wang K., Bai J. K. & Yang B. (2021). Paleogeographic evolution of a Carboniferous–Permian sea in the southernmost part of the Central Asian Orogenic Belt, NW China: Evidence from microfacies, provenance and paleobiogeography. Earth-Science Reviews, 220, 103738.

Ondrejka M., Uher P., Putiš M., Kohút M., Broska I., Larionov A., Bojar A. V. & Sobocký T. (2021). Permian A-type granites of the Western Carpathians and Transdanubian regions: products of the Pangea supercontinent breakup. International Journal of Earth Sciences, 110(6), 2133-2155.

Qiao F., Zhang Y. C., Wang Y., Yuan D. X., Ju Q., Xu H. P., Zhang H., Zheng Q. F., Cai Y. F., Hou Z. S. & Shen S. Z. (2021). An updated age of Permian strata in the Raggyorcaka and Qamdo areas, Tibet and their paleogeographic implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 582, 110660.

Tang W., Zhang Y., Pe-Piper G., Piper D. J., Guo Z. & Li W. (2021). Permian rifting processes in the NW Junggar Basin, China: Implications for the post-accretionary successor basins. Gondwana Research, 98, 107-124.

Villaseñor G., Catlos E. J., Broska I., Kohút M., Hraško Ľ., Aguilera K., Etzel T. M., Kyle J. R. & Stockli D. F. (2021). Evidence for widespread mid-Permian magmatic activity related to rifting following the Variscan orogeny (Western Carpathians). Lithos, 390, 106083.

Zhang D., Huang B., Zhao G., Meert J. G., Williams S., Zhao J., & Zhou T. (2021). Quantifying the Extent of the Paleo‐Asian Ocean During the Late Carboniferous to Early Permian. Geophysical Research Letters, 48(15), e2021GL094498.


Cui Y., Li M., Van Soelen E. E., Peterse F. & Kürschner W. M. (2021). Massive and rapid predominantly volcanic CO2 emission during the end-Permian mass extinction. Proceedings of the National Academy of Sciences, 118(37).

Davydov V. I. (2021). Tunguska сoals, Siberian sills and the Permian-Triassic extinction. Earth-Science Reviews, 212, 103438.

Fielding C. R., Frank T. D., Tevyaw A. P., Savatic K., Vajda V., McLoughlin S., Mays C. Nicoll R. S., Bocking M. & Crowley, J. L. (2021). Sedimentology of the continental end‐Permian extinction event in the Sydney Basin, eastern Australia. Sedimentology, 68(1), 30-62.

Hülse D., Lau K.V., van de Velde S.J., Arndt S., Meyer K. M. & Ridgwell A. (2021). End-Permian marine extinction due to temperature-driven nutrient recycling and euxinia. Nature Geoscience, 14, 862–867.

Liu Y., Li L., van Wijk J., Li A. & Fu Y. V. (2021). Surface-wave tomography of the Emeishan large igneous province (China): Magma storage system, hidden hotspot track, and its impact on the Capitanian mass extinction. Geology, 49(9), 1032-1037.

Luo M., Buatois L. A., Shi G. R. & Chen Z. Q. (2021). Infaunal response during the end-Permian mass extinction. GSA Bulletin, 133(1-2), 91-99.

Retallack G. J. (2021). Multiple Permian-Triassic life crises on land and at sea. Global and Planetary Change, 198, 103415.

Viglietti P. A., Benson R. B., Smith R. M., Botha J., Kammerer C. F., Skosan Z., Butler E., Crean A., Eloff B., Kaal S., Mohoi J., Molehe W., Mtalana N., Mtungata S., Ntheri N., Ntsala T., Nyaphuli J., October P., Skinner G., Strong M., Stummer H., Wolvaardt F. P. & Angielczyk K. D. (2021). Evidence from South Africa for a protracted end-Permian extinction on land. Proceedings of the National Academy of Sciences, 118(17).

Wan J., Foster W. J., Tian L., Stubbs T. L., Benton M. J., Qiu X. & Yuan A. (2021). Decoupling of morphological disparity and taxonomic diversity during the end-Permian mass extinction. Paleobiology, 47(3), 402-417.

Yang S., Wang Q., Lai X., Qiao L., Wang R., Huang Y. & Wang Y. (2021). Genesis of end-Guadalupian bauxite and pyrite deposits in the Youjiang Basin (South China): Insights into the causative link between magmatic events and mass extinction. Journal of Asian Earth Sciences, 215, 104801.

Zhang H., Zhang F., Chen J. B., Erwin D. H., Syverson D. D., Ni P., Rampino M., Chi Z., Cai Y. F., Xiang L., Li W. Q., Liu S. A., Wang R. C., Feng Z., Li H. M., Zhang T., Cai H. M., Zheng W., Cui Y., Zhu X. K., Hou Z. Q., Wu F. Y., Xu Y. G., Planavsky N. & Shen S. Z. (2021). Felsic volcanism as a factor driving the end-Permian mass extinction. Science advances, 7(47), eabh1390.

Zhu J., Zhang Z., Santosh M., Tan S. & Jin Z. (2021). Submarine basaltic eruptions across the Guadalupian-Lopingian transition in the Emeishan large igneous province: Implication for end-Guadalupian extinction of marine biota. Gondwana Research, 92, 228-238.

Vertebrate palaeontology (excluding conodonts)

Angielczyk K. D., Liu J. & Yang W. (2021). A redescription of Kunpania scopulusa, a bidentalian dicynodont (Therapsida, Anomodontia) from the? Guadalupian of northwestern China. Journal of Vertebrate Paleontology, 41(1), e1922428.

Citton P., de Valais S., Díaz-Martínez I., González S. N., Greco G. A., Cónsole-Gonella C. & Leonardi G. (2021). Age-constrained therapsid tracks from a mid-latitude upland (Permian–Triassic transition, Los Menucos Complex, Argentina). Journal of South American Earth Sciences, 110, 103367.

Gai Z., Bai Z., Lin X., Meng X. & Zhang J. (2021). First Record of Petalodus Owen, 1840 (Chondrichthyes, Petalodontidae) in the Lower Permian (Cisuralian) of China. Acta Geologica Sinica‐English Edition, 95(4), 1057-1064.

Kammerer C. F. & de los Angeles Ordoñez M. (2021). Dicynodonts (Therapsida: Anomodontia) of South America. Journal of South American Earth Sciences, 108, 103171.

Logghe A., Mujal E., Marchetti L., Nel A., Pouillon J. M., Giner S., Garrouste R. & Steyer J. S. (2021). Hyloidichnus trackways with digit and tail drag traces from the Permian of Gonfaron (Var, France): New insights on the locomotion of captorhinomorph eureptiles. Palaeogeography, Palaeoclimatology, Palaeoecology, 573, 110436.

Marsicano C., Angielczyk K. D., Cisneros J. C., Richter M., Kammerer C. F., Fröbisch J. & Smith R. M. (2021). Brazilian Permian dvinosaurs (Amphibia, Temnospondyli): revised description and phylogeny. Journal of Vertebrate Paleontology, 41(1), e1893181.

Rubidge B. S., Day M. O. & Benoit J. (2021). New specimen of the enigmatic dicynodont Lanthanostegus mohoii (Therapsida, Anomodontia) from the southwestern Karoo Basin of South Africa, and its implications for middle Permian biostratigraphy. Frontiers in Earth Science, 9, 668143.

Smith R. M., Angielczyk K. D., Benoit J. & Fernandez V. (2021). Neonate aggregation in the Permian dicynodont Diictodon (Therapsida, Anomodontia): Evidence for a reproductive function for burrows?. Palaeogeography, Palaeoclimatology, Palaeoecology, 569, 110311.


Lara-Peña R. A., Navas-Parejo P. & Torres-Martínez M. A. (2021). Permian autochthony of northwestern Mexico based on conodont paleogeographic relationships with southwestern Laurentia. Newsletters on Stratigraphy, 363-376.

Muto S., Okumura Y. & Mizuhara T. (2021). Late Kungurian conodonts of pelagic Panthalassa from seamount-capping limestone in Ogama, Kuzuu, Tochigi Prefecture, Japan. Paleontological Research, 25(2), 105-119.

Wu B., Li H., Joachimski M. M., Wignall P. B., Jiang H., Yan J., Wang L., Wu X. & Lai X. (2021). Roadian-Wordian (Middle Permian) Conodont Biostratigraphy, Sedimentary Facies and Paleotemperature Evolution at the Shuixiakou Section, Xikou Area, Southeastern Qinling Region, China. Journal of Earth Science, 32(3), 534-553.

Yuan D. X., Zhang Y. C., Qiao F., Xu H. P., Ju Q. & Shen S. Z. (2022). A new late Kungurian (Cisuralian, Permian) conodont and fusuline fauna from the South Qiangtang Block in Tibet and their implications for correlation and paleobiogeography. Palaeogeography, Palaeoclimatology, Palaeoecology, 589, 110822.

Zhang L., Wu J., Yuan D., Forel M. B., Chang S., Khan M. Z., Feng Q., He W., Ma Q., Danelian T., Caridroit M. & Ito T. (2021). Integrated Radiolarian and Conodont Biostratigraphy of the Middle to Late Permian Linghao Formation in Northwestern Guangxi, South China. Acta Geologica Sinica‐English Edition, 95(6), 1984-1997.

Foraminifera, algae and microbes

Arefifard S. & Clapham M. E. (2021). Cisuralian and Guadalupian global paleobiogeography of fusulinids in response to tectonics, ocean circulation and climate change. Palaeogeography, Palaeoclimatology, Palaeoecology, 561, 110052.

Ju Q., Zhang Y. C., Qiao F. & Xu H. P. (2021). A Middle Permian assemblage of smaller foraminifera (Shanita-Hemigordiopsis assemblage) from the central Lhasa Block and its paleobiogeographic implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 572, 110417.

Mays C., McLoughlin S., Frank T. D., Fielding C. R., Slater S. M. & Vajda V. (2021). Lethal microbial blooms delayed freshwater ecosystem recovery following the end-Permian extinction. Nature communications, 12(1), 1-11.

Mays, C. Vajda V. & McLoughlin S. (2021). Permian–Triassic non-marine algae of Gondwana—Distributions, natural affinities and ecological implications. Earth-Science Reviews, 212, 103382.

Nestell M. K. & Nestell G. P. (2021). An early Capitanian (Middle Permian) foraminiferal connection between southern British Columbia, Canada and South Primorye, Russian Far East. PalZ, 1-22.

Sharahi S. Y., Yeganeh B. Y., Arefifard S., Vachard D. & Farahpour M. M. (2021). Biostratigraphy, taxonomy and paleobiogeography of the upper Cisuralian (upper Yakhtashian–Bolorian) foraminifers from east-central Iran, with clarification of the taxonomy of the fusulinid genera Cuniculinella and Cuniculina pre-occupied. Journal of Paleontology, 95(S81), 1-30.

Su C., Chen Z. Q., Wang X., Wu S. & Guo Z. (2021). Biotic and palaeoecological variations in the Permian-Triassic boundary microbialite (Xiejiacao, South China): Implication for a two-phase ecological crisis in microbialite ecosystems. Global and Planetary Change, 207, 103679.

Ueno K. (2021). Thailandina and Neothailandina and their family Thailandinidae salvaged: a valid taxonomic group of peculiar Permian fusuline Foraminifera. Journal of Paleontology, 1-6.

Brachiopods, bivalves and other invertebrates

Afanasjeva G. A. & Viskova L. A. (2021). Morphophysiological Peculiarities of Articulated Brachiopods and Marine Bryozoans as a Reason for Their Different Evolutionary Consequences of the Permian–Triassic Crisis. Paleontological Journal, 55(7), 742-751.

Asato K. & Kase T. (2021). Gigantic scaphopods (Mollusca) from the Permian Akasaka Limestone, central Japan. Journal of Paleontology, 95(4), 748-762.

Baud A., Richoz S., Brandner R., Krystyn L., Heindel K., Mohtat T., Mohtat-Agai P. & Horacek M. (2021). Sponge takeover from end-Permian mass extinction to Early Induan time: Records in Central Iran microbial buildups. Frontiers in Earth Science, 9.

Biakov A. S. & Kutygin R. V. (2021). First Record of the Genus Unionites Wissmann (Bivalvia) in the Boreal Permian of Northeast Russia. Paleontological Journal, 55(4), 372-377.

Biakov A. S., Danukalova M. K., Kuzmichev A. B. & Sobolev E. S. (2021). Permian Bivalves of the Pronchishchev Ridge (North Siberia): New Data on Taxonomic Composition, Biostratigraphy, and Biogeographic Relationships. Paleontological Journal, 55(3), 259-271.

Chen F., Xue W., Yan J. & Meng Q. (2021). The implications of the giant bivalve family Alatoconchidae for the end‐Guadalupian (Middle Permian) extinction event. Geological Journal, 56(12), 6073-6087.

Chitnarin A. & Ketwetsuriya C. (2021, October). Middle Permian ostracod fauna from the Khao Khad Formation (Indochina Terrane), Central Thailand. Annales de Paléontologie, 107(4), 102521.

Guo Z., Chen Z. Q., Harper D. A. & Huang Y. (2021). Permian–Triassic phylogenetic and morphologic evolution of rhynchonellide brachiopods. Paleobiology, 1-21.

Ketwetsuriya C., Hausmann I. M. & Nützel A. (2021). Diversity patterns of Middle Permian gastropod assemblages from the Tak Fa Formation, Central Thailand. Palaeobiodiversity and Palaeoenvironments, 101(4), 907-925.

Logghe A., Nel A., Steyer J. S., Ngô-Muller V., Pouillon J. M. & Garrouste R. (2021). A twig-like insect stuck in the Permian mud indicates early origin of an ecological strategy in Hexapoda evolution. Scientific reports, 11(1), 1-7.

Masunaga M. & Shiino Y. (2021). Death or living assemblage? The middle Permian discinid brachiopods in the Kamiyasse area, Southern Kitakami Mountains, northeastern Japan. Paleontological Research, 25(3), 258-278.

Naugolnykh S. V. & Bicknell R. D. (2021). Ecology, morphology and ontogeny of Paleolimulus kunguricus—a horseshoe crab from the Kungurian (Cisuralian) of the Cis‐Urals, Russia. Lethaia.

Niko S. & Badpa M. (2021). Tabulate corals from the Middle and Upper Permian formations in the Julfa area, Northwest Iran. Bulletin of the National Museum of Nature and Science. Series C (Geology & Paleontology), 47, 41-51.

Pretorius A. I., Labandeira C. C., Nel A. & Prevec R. (2021). Latest Permian insects from Wapadsberg Pass, southern Karoo Basin, South Africa. Austral Entomology, 60(3), 560-570.

Quiroz-Barroso S. A., Torres-Martínez M. A. & Sour-Tovar F. (2021). Gastropods from the Paso Hondo Formation (lower-middle Permian) of Chiapas, Mexico: Stratigraphical and paleobiogeographical considerations. Journal of South American Earth Sciences, 110, 103350.

Tarnac A., Forel M. B., Nestell G., Nestell M. & Crasquin S. (2021). Middle Permian ostracods (Crustacea) from the Guadalupe Mountains, West Texas, USA. European Journal of Taxonomy, 770, 1-60.

Viaretti M., Crippa G., Posenato R., Shen S. Z. & Angiolini L. (2021). Lopingian brachiopods from the Abadeh section (Central Iran) and their biostratigraphic implications. Bollettino della Società Paleontologica Italiana, 60(3), 213-254.

Wang X., Yao L. & Wang X. (2021). Permian naotic‐dissepimented rugose corals in China and their palaeoenvironmental implications. Geological Journal, 56(12), 6151-6161.

Wu H. T., Zhang Y. & Sun Y. L. (2021). A brachiopod fauna from latest Permian to Induan of northern Guizhou, South China and its evolutionary pattern. Geological Journal, 56(12), 6189-6198.

Xu H. P., Aung K. P., Zhang Y. C., Shi G. R., Cai F. L., Zaw T., Ding L., Sein K. & Shen S. Z. (2021). A late Cisuralian (Early Permian) brachiopod fauna from the Taungnyo Group in the Zwekabin Range, eastern Myanmar and its biostratigraphic, paleobiogeographic, and tectonic implications. Journal of Paleontology, 95(6), 1158-1188.

Yang A., Huang D., Clark B. & Sendino C. (2021). Description of new Permian orthocerid forms from Gufeng Formation of South China (the Yangtze Craton) and their palaeobiogeographic implications in the Palaeotethys. Historical Biology, 33(11), 2804-2821.

Zhang Y. (2021). Shell microstructures of latest Permian Rugosochonetidae (Brachiopoda): evidence from SEM‐and CT‐scanned shell materials. Lethaia.

Palaeobotany and Palynology

Blomenkemper P., Baeumer R., Backer M., Abu Hamad A., Wang J., Kerp H. & Bomfleur B. (2021). Bennettitalean Leaves From the Permian of Equatorial Pangea—The Early Radiation of an Iconic Mesozoic Gymnosperm Group. Frontiers in Earth Science, 9, 162.

Bodnar J., Coturel E. P., Falco J. I. & Beltrán M. (2021). An updated scenario for the end-Permian crisis and the recovery of Triassic land flora in Argentina. Historical Biology, 33(12), 3654-3672.

Cai Y. F., Zhang H., Cao C. Q., Zheng Q. F., Jin C. F. & Shen S. Z. (2021). Wildfires and deforestation during the Permian–Triassic transition in the southern Junggar Basin, Northwest China. Earth-Science Reviews, 218, 103670.

Cai Y. F., Zhang H., Feng Z. & Shen S. Z. (2021). Intensive wildfire associated with volcanism promoted the vegetation changeover in Southwest China during the Permian− Triassic transition. Frontiers in Earth Science, 58.

Cariglino B., Moisan P. & Lara M. B. (2021). The fossil record of plant-insect interactions and associated entomofaunas in Permian and Triassic floras from southwestern Gondwana: A review and future prospects. Journal of South American Earth Sciences, 111, 103512.

Fernández J. A. (2021). Revision of Botrychiopsis plantiana: A Key Species of the Gzhelian–Cisuralian in Westernmost Gondwana. Ameghiniana, 58(1), 1-11.

Gibson M. E. & Bodman D. J. (2021). Evaporite palynology: a case study of the Permian (Lopingian) Zechstein Sea. Journal of the Geological Society, 178(6).

Gibson M. E. & Wellman C. H. (2021). The use of spore–pollen assemblages to reconstruct vegetation changes in the Permian (Lopingian) Zechstein deposits of northeast England. Review of Palaeobotany and Palynology, 288, 104399.

Guo Y., Zhou Y., Bek J., Yang S. L. & Feng Z. (2021). Qasimia yunnanica sp. nov., a marattialean fern with bivalvate synangia from the Lopingian of Southwest China. Review of Palaeobotany and Palynology, 293, 104497.

Koll R. A. & DiMichele W. A. (2021). Dominance-diversity architecture of a mixed hygromorphic-to-xeromorphic flora from a botanically rich locality in western equatorial Pangea (lower Permian Emily Irish site, Texas, USA. Palaeogeography, Palaeoclimatology, Palaeoecology, 563, 110132.

Lopes G., Pereira Z., Fernandes P., Marques J., Mendes M. & Götz A. E. (2021). Permian stratigraphy and palynology of the Lower Karoo Group in Mozambique–a 2020 perspective. Newsletters on Stratigraphy.

Mangerud G., Paterson N. W. & Bujak J. (2021). Permian palynoevents in the circum-Arctic region. Atlantic Geology: Journal of the Atlantic Geoscience Society/Atlantic Geology: revue de la Société Géoscientifique de l’Atlantique, 57, 57-69.

Pšenička J., Wang J., Bek J., Pfefferkorn H. W., Opluštil S., Zhou W., Frojdová J. V. & Libertin M. (2021). A zygopterid fern with fertile and vegetative parts in anatomical and compression preservation from the earliest Permian of Inner Mongolia, China. Review of Palaeobotany and Palynology, 294, 104382.

Stephenson M. H. & Korngreen D. (2021). Palynology of the Permian of the Makhtesh Qatan-2, Ramon-1 and Boqer-1 boreholes Arqov Formation, Negev, Israel, Rivista Italiana di Paleontologia e Stratigrafia, 127(3), 655-672.

Wan M., Yang W., Wang K., Liu L. & Wang J. (2021). Zhuotingoxylon liaoi gen. et sp. nov., a silicified coniferous trunk from the Changhsingian (Permian) of southern Bogda Mountains, northwestern China. Geological Journal, 56(12), 6135-6150.

Wu Q., Ramezani J., Zhang H., Wang J., Zeng F., Zhang Y., Liu F., Chen J., Cai Y., Hou Z., Liu C., Yang W., Henderson C. M. & Shen S. Z. (2021). High-precision U-Pb age constraints on the Permian floral turnovers, paleoclimate change, and tectonics of the North China block. Geology, 49(6), 677-681.

Xiong C., Wang J., Huang P., Cascales-Minana B., Cleal C. J., Benton M. J. & Xue J. (2021). Plant resilience and extinctions through the Permian to Middle Triassic on the North China Block: A multilevel diversity analysis of macrofossil records. Earth-Science Reviews, 223, 103846.

Zhou Y., Guo Y., Pšenička J., Bek J., Yang S. L. & Feng Z. (2021). A new marattialean fern, Pectinangium xuanweiense sp. nov., from the Lopingian of Southwest China. Review of Palaeobotany and Palynology, 295, 104500.

Zhou W., Pšenička J., Bek J., Wan M., Boyce C. K. & Wang J. (2021). A new anachoropterid fern from the Asselian (Cisuralian) Wuda Tuff Flora. Review of Palaeobotany and Palynology, 294, 104346.

Zhou W. M., Pšenička J., Hilton J. & Wang J. (2021). Palaeobiogeographical implications of the earliest botryopterid ferns in Cathaysia. Historical Biology, 33(11), 2577-2583.


Feng Q., Qiu N., Fu X., Li W., Xu Q., Li X. & Wang J. (2021). Permian geothermal units in the Sichuan Basin: Implications for the thermal effect of the Emeishan mantle plume. Marine and Petroleum Geology, 132, 105226.

Hou L., Ma W., Luo X., Liu J., Liu S. & Zhao Z. (2021). Hydrocarbon generation-retention-expulsion mechanism and shale oil producibility of the Permian Lucaogou shale in the Junggar Basin as simulated by semi-open pyrolysis experiments. Marine and Petroleum Geology, 125, 104880.

Irakulis-Loitxate I., Guanter L., Liu Y. N., Varon D. J., Maasakkers J. D., Zhang Y., Chulakadabba I., Wofsy S. C., Thorpe A. K., Duren R. M., Frankenberg C., Lyon D. R., Hmiel B., Cusworth D. H., Zhang Y., Segl K., Gorroño J., Sánchez-García E., Sulprizio M. P., Cao K., Zhu H., Liang J., Li X., Aben I. & Jacob D. J. (2021). Satellite-based survey of extreme methane emissions in the Permian basin. Science Advances, 7(27), eabf4507.

Li X., Dai S., Nechaev V. P., Graham I. T., French D., Wang X., Zhao L. & Zhao J. (2021). Mineral matter in the late Permian C1 coal from Yunnan Province, China, with emphasis on its origins and modes of occurrence. Minerals, 11(1), 19.

Li Y., Fan A., Yang R., Sun Y. & Lenhardt N. (2021). Sedimentary facies control on sandstone reservoir properties: A case study from the Permian Shanxi Formation in the southern Ordos basin, central China. Marine and Petroleum Geology, 129, 105083.

Li J., Zhang X., Tian J., Liang Q. & Cao T. (2021). Effects of deposition and diagenesis on sandstone reservoir quality: A case study of Permian sandstones formed in a braided river sedimentary system, northern Ordos Basin, Northern China. Journal of Asian Earth Sciences, 213, 104745.

Lin M., Xi K., Cao Y., Liu Q., Zhang Z. & Li K. (2021). Petrographic features and diagenetic alteration in the shale strata of the Permian Lucaogou Formation, Jimusar Sag, Junggar Basin. Journal of Petroleum Science and Engineering, 203, 108684.

Liu J., Dai S., Song H., Nechaev V. P., French D., Spiro B. F., Graham I. T., Hower J. C., Shao L. & Zhao J. (2021). Geological factors controlling variations in the mineralogical and elemental compositions of Late Permian coals from the Zhijin-Nayong Coalfield, western Guizhou, China. International Journal of Coal Geology, 247, 103855.

Ling K., Wen H., Zhang Q., Luo C., Gu H., Du S. & Yu W. (2021). Super-enrichment of lithium and niobium in the upper Permian Heshan Formation in Pingguo, Guangxi, China. Science China Earth Sciences, 64(5), 753-772.

Pei Y., Yu W., Sepehrnoori K., Gong Y., Xie H. & Wu K. (2021). The influence of development target depletion on stress evolution and infill drilling of upside target in the Permian Basin. SPE Reservoir Evaluation & Engineering, 24(03), 570-589.

Shen M., Dai S., Graham I. T., Nechaev V. P., French D., Zhao F., Shao L., Liu S., Zuo J., Zhao J., Chen K. & Xie X. (2021). Mineralogical and geochemical characteristics of altered volcanic ashes (tonsteins and K-bentonites) from the latest Permian coal-bearing strata of western Guizhou Province, southwestern China. International Journal of Coal Geology, 237, 103707.

Singh D. P., Hazra B., Wood D. A. & Singh P. K. (2021). Hydrocarbon generation and kinetics: A case study of Permian shales, India. Journal of Asian Earth Sciences, 222, 104960.

Spina A., Cirilli S., Sorci A., Schito A., Clayton G., Corrado S., Fernandes P., Galasso F., Montesi G., Pereira Z., et al. 2021. Assessing Thermal Maturity through a Multi-Proxy Approach: A Case Study from the Permian Faraghan Formation (Zagros Basin, Southwest Iran). Geosciences, 11, 484.

Yang Y., Yang Y., Wen L., Zhang X., Chen C., Chen K., Zhang Y., Di G., Wang H. & Xie C. (2021). New progress and prospect of Middle Permian natural gas exploration in the Sichuan Basin. Natural Gas Industry B, 8(1), 35-47.

Yu W., Polgári M., Fintor K., Gyollai I., Szabó M., Velledits F., Liu Z. & Du Y. (2021). Contribution of microbial processes to the enrichment of Middle Permian manganese deposits in northern Guizhou, South China. Ore Geology Reviews, 136, 104259.

Zhi D, Tang Y., He W., Guo X., Zheng M. & Huang L. (2021). Orderly coexistence and accumulation models of conventional and unconventional hydrocarbons in Lower Permian Fengcheng Formation, Mahu sag, Junggar Basin. Petroleum Exploration and Development, 48(1), 43-59.